The 386 microcode sequencer has a one-cycle pipeline delay: when a jump or RNI (run next instruction) is decoded, the micro-instruction immediately after it has already been fetched and will execute before the jump takes effect. This "delay slot" is a basic property of the sequencer, and the microcode is written to fill it with useful work rather than waste a cycle on a bubble. The examples in the PTSAV section above show this: at 582/5AE, the micro-instruction after LCALL executes before the subroutine begins.
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Implementations have found ways to optimize transform pipelines by collapsing identity transforms, short-circuiting non-observable paths, deferring buffer allocation, or falling back to native code that does not run JavaScript at all. Deno, Bun, and Cloudflare Workers have all successfully implemented "native path" optimizations that can help eliminate much of the overhead, and Vercel's recent fast-webstreams research is working on similar optimizations for Node.js. But the optimizations themselves add significant complexity and still can't fully escape the inherently push-oriented model that TransformStream uses.